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The predominantly African origin of all modern human populations is well established, but the route taken out of Africa is still unclear. Two alternative routes, via Egypt and Sinai or across the Bab el Mandeb strait into Arabia, have traditionally been proposed as feasible gateways in light of geographic, paleoclimatic, archaeological, and genetic evidence. Distinguishing among these alternatives has been difficult. West Eurasian components were masked out, and the remaining African haplotypes were compared with a panel of sub-Saharan African and non-African genomes.
We showed that masked Northeast African haplotypes overall were more similar to non-African haplotypes and more frequently present outside Africa than were any sets of haplotypes derived from a West African population. Furthermore, the masked Egyptian haplotypes showed these properties more markedly than the masked Ethiopian haplotypes, pointing to Egypt as the more likely gateway in the exodus to the rest of the world. Using five Ethiopian and three Egyptian high-coverage masked genomes and the multiple sequentially Markovian coalescent MSMC approach, we estimated the genetic split times of Egyptians and Ethiopians from non-African populations at 55, and 65, years ago, respectively, whereas that of West Africans was estimated to be 75, years ago.
The routes followed by fully modern humans as they expanded out of Africa 50,—, years ago into Eurasia have long been a central question of anthropology 1 and have important implications for understanding the evolutionary history of all non-African populations. So far, neither fossil and archaeological 2—4 nor genetic 5,6 evidence has been able to distinguish between an exit through Egypt and Sinai northern route 7 or one through Ethiopia, the Bab el Mandeb strait, and the Arabian Peninsula southern route.
Information to discriminate between the northern and southern routes might still be present in Africa within the full genomes of the populations inhabiting modern Egypt and the Horn of Africa, and thus further investigation is warranted. However, although it might not be easy to extract this information because of the past and recent genetic introgression experienced by these populations, 13,14 full sequences of Northeast African genomes would provide the best starting point for these and other analyses.
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